Various other wider-scale studies out of archosaur relationships by Nesbitt et al

Various other wider-scale studies out of archosaur relationships by Nesbitt et al


None Parker et al. ( 2005 ) nor Take a look mais aussi al. ( 2005 ) tried a great phylogenetic investigation out of Roentgen. callenderi in line with the the fresh skeletal situation. Yet not, each other organizations detailed multiple popular features of the new hindlimb which were unambiguous synapomorphies of Pseudosuchia. Parker mais aussi al. ( 2005 ) hypothesized a prospective sibling-group experience of aetosaurs, detailing the brand new morphological parallels of squamosal and you will paramedian osteoderms. An identical position are recommended of the Hunt ainsi que al. ( 2005 ) predicated on unspecified character states of the tarsus and the osteoderms. Brusatte mais aussi al. ( 2010 ) incorporated Roentgen. callenderi within their study regarding early archosaur relationships, programming regarding authored numbers and you may meanings of Parker mais aussi al. ( 2005 ) and you may Seem et al. ( 2005 ). One to analysis retrieved R. callenderi because brother taxon on early-diverging suchian clade Ornithosuchidae. ( 2011 ) retrieved R. callenderi in a properly-served updates as sibling taxon so you can aetosaurs, a consequence supported by newer training (elizabeth.g., Butler mais aussi al., 2014 ; ; Nesbitt & Butler, 2013 ; Nesbitt ainsi que al., 2014 ; Nesbitt et al., 2020 ; Nesbitt, Stocker, mais aussi al., 2018 ).

This sum totally describes the newest osteology regarding R. callenderi based on at least a dozen anyone (considering femora amount) off PEFO in addition to about three well-maintained relevant limited skeletons gathered ranging from 2005 and 2015, brings osteohistological perspective, and you may discusses this new taxon’s advantages to possess wisdom early suchian development.

dos.1 Stratigraphic standing

That it breakdown away from Roentgen. callenderi was mostly according to issue regarding several localities on the Chinle Development out of Petrified Tree Federal Playground (PEFO), Arizona, Us (Shape 1; get a hold of Systematic Paleontology). The top of Triassic Chinle Development in this field comprises mostly fluvial and you can floodplain strata off alternating claystones, siltstones, sandstones, and you can conglomerates (Billingsley, 1985 ; Heckert & Lucas, 2002a ; ). The latest Chinle Creation regarding Petrified Forest Federal Playground urban area saves well-recognized traditional assemblages of plant life (palynomorphs, will leave, wood), invertebrates, and you may vertebrates (age.grams., Ash, 2005 ; Baranyi ainsi que al., 2018 ; An excellent, 1998 ; Irmis, 2005 ; Litwin, Navigate, & Ash, 1991 ; Parker, 2005b , 2006 ; Reichgelt et al., 2013 ). Very non-dentigerous and you can non-osteoderm specimens off R. callenderi regarding PEFO, together with those people demonstrated here, have been discovered quickly a lot more than Decorated Wilderness Sandstone step three (sensu Billingsley, 1985 ; Lithodendron Wash Sleep sensu Heckert & Lucas, 2002a ) in between part of the Petrified Tree Affiliate (sensu Woody, 2006 ) of one’s upper a portion of the Chinle Development (Heckert & Lucas, 2002a ; Irmis, 2005 ; Padian, 1990 ; Parker, 2005b , 2006 ; Parker et al., 2005 ; Parker & ). Separated pearly whites from Roentgen. callenderi was in fact recovered regarding microvertebrate assemblages away from lower in the fresh new Chinle Development within PEFO, extending the latest stratigraphic selection of so it taxon into the Sonsela User and uppermost Blue Mesa Member (find Medical Paleontology).

The 3 fundamental localities you to definitely brought the fresh new specimens out-of Roentgen. callenderi revealed here (Studies S1) may be the Revueltosaurus Quarry (PFV 297), Revueltosaurus Quarry SW (PFV 414), and the Providing Website (PFV 231). The internet sites exists as paleosols in the a stacked fluvial succession when you look at the the middle of the newest Petrified Forest Associate (Atchley et al., 2013 ; Loughney ainsi que al., 2011 ; Parker et al., 2005 ; Parker & ). PFV 297 and you can PFV 414 are separated spatially from the 20 yards and generally are in turn about 200 yards to the southeast of PFV 231 (Figure 1a). Coated Desert Sandstone step three, hence quickly underlies every three localities, was stratigraphically

40–fifty yards over the base of the Petrified Tree Representative (Figure 1b; Parker & ), and you may signifies a sheet sandstone deposited from the recurrent move into the a high sinuosity lake route (Johns, 1988 ). PFV 297 and you can PFV 414 can be found quite high regarding sequence than just PFV 231 (Parker & ).

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